<div dir="ltr"><div class="gmail_quote"><div dir="ltr"><div style="font-family:tahoma,sans-serif"><br><table cellspacing="0" cellpadding="0" width="100%" bgcolor="#474444" style="font-family:Arial,Helvetica,sans-serif"><tbody><tr><td style="font-size:0px;height:33px;line-height:0"></td></tr><tr><td valign="top"><table cellspacing="0" cellpadding="0" width="600" align="center" border="0" style="width:600px;margin:0px auto"><tbody><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:0px;text-align:left;background-color:rgb(254,251,251)"><table cellspacing="0" cellpadding="0" align="left" style="width:600px"><tbody><tr height="44" style="height:44px"><th style="border-width:medium;border-style:none;border-color:initial;width:477px;vertical-align:middle;font-weight:normal;padding:10px 0px 0px 10px;text-align:left;background-color:rgb(66,176,91)"><p align="left" style="margin-bottom:1em;font-size:14px;font-family:tahoma,geneva,sans-serif;color:rgb(253,251,251);margin-top:0px;line-height:18px;background-color:transparent"><strong> Seminario IFEVA</strong></p></th><th style="border-width:medium;border-style:none;border-color:initial;width:103px;vertical-align:middle;font-weight:normal;padding:0px 5px;text-align:left;background-color:transparent"><div style="padding:10px;text-align:right"><table cellspacing="0" cellpadding="0" border="0" style="display:inline-block"><tbody><tr><td style="padding-right:5px"><a href="https://twitter.com/ifevaok?ref_src=twsrc%5Etfw%7Ctwcamp%5Eembeddedtimeline%7Ctwterm%5Eprofile%3Aifevaok&ref_url=http%3A%2F%2Fwww.ifeva.edu.ar%2Findex.tw.html" target="_blank"><img title="Twitter" alt="Twitter" src="https://mailchef.s3.amazonaws.com/uploads/mailstyler/images/478A2A14-0536-35C1-A7E4-6A014235947D_Image_1_9f1117f4-4654-4d62-878f-885c62b8fa59.png" width="24" style="border-width:medium;border-style:none;border-color:initial;display:block"></a></td><td style="padding-right:5px"><a href="https://www.youtube.com/channel/UCCC_QmqL9yuKBnZ6p4mnxBw" target="_blank"><img title="Youtube" alt="Youtube" src="https://mailchef.s3.amazonaws.com/uploads/mailstyler/images/8EF10070-9897-5A56-C849-AF5EF89AEA87_Image_2_e946f5fa-93af-4d25-972f-4211dca72430.png" width="24" style="border-width:medium;border-style:none;border-color:initial;display:block"></a></td><td><a href="https://www.instagram.com/ifevaok/" target="_blank"><img title="Instagram" alt="Instagram" src="https://mailchef.s3.amazonaws.com/uploads/mailstyler/images/371C7FF9-A40B-509F-8253-F6AC579686D2_Image_3_79b8ac76-96a9-425d-adfc-620fc1609bf9.png" width="24" style="border-width:medium;border-style:none;border-color:initial;display:block"></a></td></tr></tbody></table></div></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:580px;font-weight:normal;padding:0px 20px 0px 0px;text-align:left;background-color:rgb(254,251,251)"><table cellspacing="0" cellpadding="0" align="left" style="width:580px"><tbody><tr height="92" style="height:92px"><th style="border-width:medium;border-style:none;border-color:initial;width:580px;vertical-align:middle;font-weight:normal;padding:0px;text-align:left;background-color:transparent"><table cellspacing="0" cellpadding="0" align="center" border="0" style="margin:0px auto"><tbody><tr><td align="center" style="padding:0px"><table cellspacing="0" cellpadding="0" border="0"><tbody><tr><td style="border-width:medium;border-style:none;border-color:initial;background-color:transparent"><img alt="" src="https://mailchef.s3.amazonaws.com/uploads/mailstyler/images/152EFEE1-9961-67DB-42C9-BDFED3ECD20D_Image_4_2f0431fb-1968-435e-bed4-e7fa8fc21754.png" width="570" hspace="0" vspace="0" style="border-width:medium;border-style:none;border-color:initial;display:block"></td></tr></tbody></table></td></tr></tbody></table></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:0px;text-align:left;background-color:rgb(254,251,251)"><table cellspacing="0" cellpadding="0" align="left" style="width:600px"><tbody><tr height="76" style="height:76px"><th style="border-width:medium;border-style:none;border-color:initial;width:326px;vertical-align:middle;font-weight:normal;padding:10px 10px 0px 20px;text-align:left;background-color:rgb(243,54,62)"><p align="left" style="margin-bottom:1em;font-size:14px;color:rgb(253,251,251);margin-top:0px;line-height:18px;background-color:transparent"><strong><font face="tahoma, geneva, sans-serif">MIERCOLES</font><br><font face="tahoma, sans-serif">19</font><font face="tahoma, geneva, sans-serif"> de Octubre</font></strong><strong style="background-color:transparent"><font face="tahoma, geneva, sans-serif"> </font><font face="tahoma, sans-serif">2</font><font face="tahoma, geneva, sans-serif">022</font></strong></p><p align="left" style="margin-bottom:1em;font-size:14px;color:rgb(253,251,251);margin-top:0px;line-height:18px;background-color:transparent"><strong><font face="tahoma, geneva, sans-serif">12.00 hs</font></strong></p></th><th style="border-width:medium;border-style:none;border-color:initial;width:244px;vertical-align:middle;padding:10px 0px 0px;text-align:left;background-color:rgb(108,108,108)"><p align="center" style="font-weight:normal;margin-bottom:1em;font-size:14px;font-family:tahoma,geneva,sans-serif;color:rgb(56,56,56);text-align:center;margin-top:0px;line-height:18px;background-color:transparent"><br></p><p align="center" style="font-weight:normal;margin-bottom:1em;font-size:14px;font-family:tahoma,geneva,sans-serif;color:rgb(56,56,56);text-align:center;margin-top:0px;line-height:18px;background-color:transparent"><a title="" href="https://www.youtube.com/channel/UCCC_QmqL9yuKBnZ6p4mnxBw" style="color:rgb(253,251,251);text-decoration-line:none" target="_blank"><font color="#fdfbfb">Virtual: </font><strong>YouTube</strong><font color="#fdfbfb"> Canal </font><strong>IFEVA</strong></a></p><p align="center" style="margin-bottom:1em;font-size:14px;font-family:tahoma,geneva,sans-serif;text-align:center;margin-top:0px;line-height:18px;background-color:transparent"><font color="#fff2cc"><span style="font-weight:normal">Presencial: </span>S.U.M.<font style="font-weight:normal"> </font></font><strong><font color="#fff2cc">IFEVA</font></strong><br></p></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:0px;text-align:left;background-color:rgb(254,251,251)"><table cellspacing="0" cellpadding="0" align="left" style="width:600px"><tbody><tr height="121" style="height:121px"><th style="border-width:medium;border-style:none;border-color:initial;width:600px;vertical-align:middle;font-weight:normal;padding:0px;text-align:left;background-color:transparent"><table cellspacing="0" cellpadding="0" align="center" border="0" style="margin:0px auto"><tbody><tr><td align="center" style="padding:0px"><table cellspacing="0" cellpadding="0" border="0"><tbody><tr><td style="border-width:medium;border-style:none;border-color:initial;background-color:transparent"><img alt="" src="https://mailchef.s3.amazonaws.com/uploads/mailstyler/images/A1D4E880-2191-4DD7-2282-E4EBD8D61287_Image_5_91564d99-8f65-4b8d-9d26-1320ac7ed277.jpg" width="600" hspace="0" vspace="0" style="border-width:medium;border-style:none;border-color:initial;display:block"></td></tr></tbody></table></td></tr></tbody></table></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:10px 0px 5px;text-align:left;background-color:rgb(253,251,251)"><table cellspacing="0" cellpadding="0" align="left" style="width:600px"><tbody><tr height="126" style="height:126px"><th style="border-width:medium;border-style:none;border-color:initial;width:54px;vertical-align:top;font-weight:normal;padding:10px 5px 5px;text-align:left;background-color:transparent"><span class="gmail_default" style="font-family:tahoma,sans-serif"></span><table cellspacing="0" cellpadding="0" align="center" border="0" style="margin:0px auto"><tbody><tr><td align="center" style="padding:0px"><table cellspacing="0" cellpadding="0" border="0"><tbody><tr><td style="border-width:medium;border-style:none;border-color:initial;background-color:transparent"><img alt="" src="https://mailchef.s3.amazonaws.com/uploads/mailstyler/images/5A4A5834-3F9A-6A31-826B-CB9282750DB7_Image_6_f652f2ce-f80b-4f92-bd91-eb57b875ac83.png" width="18" hspace="0" vspace="0" style="border-width:medium;border-style:none;border-color:initial;display:block"></td></tr></tbody></table></td></tr></tbody></table></th><th style="border-width:medium;border-style:none;border-color:initial;width:526px;vertical-align:top;padding:5px"><b style="font-family:Tahoma,sans-serif"><font size="6">Photoperiodic
regulation of wheat heading time</font></b><font size="6">.<br></font></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:5px 0px;text-align:left;background-color:rgb(254,251,251)"><table cellspacing="0" cellpadding="0" align="left" style="width:600px"><tbody><tr height="564" style="height:564px"><th style="border-width:medium;border-style:none;border-color:initial;width:570px;vertical-align:top;padding:10px 15px;text-align:left;background-color:transparent"><p align="left" style="margin-bottom:1em;margin-top:0px;line-height:20px;background-color:transparent"><font size="4" style="font-family:tahoma,geneva,sans-serif"><span style="color:rgb(66,176,91)">Autor<span class="gmail_default" style="font-family:tahoma,sans-serif">es</span>:</span><font color="#383838"><span style="font-weight:normal"> </span></font></font><font size="4"><span style="font-family:Tahoma,sans-serif;font-weight:normal;background-color:transparent">Maria
Alejandra Alvarez, Chengxia Li, Huiqiong Lin, Anna Joe, Daniel Woods y </span><span style="font-family:Tahoma,sans-serif;background-color:transparent"><i>Jorge Dubcovsky(*)</i></span></font></p><p align="left" style="margin-bottom:1em;margin-top:0px;line-height:20px;background-color:transparent"><font size="4"><span style="font-family:Tahoma,sans-serif;background-color:transparent"><i>(*)</i><span style="font-weight:normal"> expositor</span></span></font></p><p align="left" style="margin-bottom:1em;margin-top:0px;line-height:20px;background-color:transparent"><strong style="background-color:transparent;color:rgb(56,56,56)"><span style="color:rgb(66,176,91)"><font face="tahoma, geneva, sans-serif" style="font-size:large">Filiaci</font><font face="tahoma, sans-serif" style="font-size:large">ó</font><font face="tahoma, geneva, sans-serif" style="font-size:large">n</font><font face="tahoma, geneva, sans-serif"><font size="4">: </font></font></span></strong><span style="background-color:transparent;font-family:Tahoma,sans-serif;font-weight:normal"><font size="4">University
of California Davis / Howard Hughes Medical Institute</font></span></p><p align="left" style="margin-bottom:1em;margin-top:0px;line-height:20px;background-color:transparent"><font size="4" style="background-color:transparent;font-weight:normal"><strong><span style="color:rgb(66,176,91)"><font face="tahoma, geneva, sans-serif">Resumen: </font></span></strong></font><span style="background-color:transparent;font-weight:normal"><font size="4">The photoperiodic response is critical for plants to adjust their reproductive phase to the most favorable season. Wheat heads earlier under long days (LD) than under short days (SD) and this difference is mainly regulated by the PHOTOPERIOD1 (PPD1) gene. Tetraploid wheat plants carrying the Ppd-A1a allele with a large deletion in the promoter head earlier under SD than plants carrying the wildtype Ppd-A1b allele with an intact promoter. Phytochromes PHYB and PHYC are necessary for the light activation of PPD1, and mutations in either of these genes result in the downregulation of PPD1 and very late heading time. We show here that both effects are reverted when the phyB mutant is combined with loss-of-function mutations in EARLY FLOWERING 3 (ELF3), a component of the Evening Complex (EC) in the circadian clock. We also show that the wheat ELF3 protein interacts with PHYB and PHYC, is rapidly modified by light, and binds to the PPD1 promoter in planta (likely as part of the EC). Deletion of the ELF3 binding region in the Ppd-A1a promoter results in its upregulation at dawn, similar to PPD1 alleles with intact promoters in the elf3 mutant background. The upregulation of PPD1 is correlated with the upregulation of the florigen gene FLOWERING LOCUS T1 (FT1) and early heading time. Loss-of-function mutations in ppd1 result in the downregulation of FT1 and delayed heading, even when combined with the elf3 mutation. Taken together, these results indicate that ELF3 operates downstream of PHYB as a direct transcriptional repressor of PPD1, and that this repression is relaxed both by light and by the deletion of the ELF3 binding region in the Ppd-A1a promoter. In summary, the regulation of the light mediated activation of PPD1 by ELF3 is critical for the photoperiodic regulation of wheat heading time.</font></span></p><p align="left" style="margin-bottom:1em;margin-top:0px;line-height:20px;background-color:transparent"><font size="4" style="background-color:transparent;color:rgb(56,56,56);font-family:tahoma,geneva,sans-serif;font-weight:normal"><strong><span style="color:rgb(66,176,91)">Palabras claves: </span></strong></font><span style="background-color:transparent;font-weight:normal"><font size="4">Early flowering, Photoperiod, Wheat</font></span><br></p><p align="left" style="font-weight:normal;margin-bottom:1em;font-family:tahoma,geneva,sans-serif;color:rgb(56,56,56);margin-top:0px;line-height:20px;background-color:transparent"></p></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:5px 0px;text-align:left;background-color:rgb(253,251,251)"><table cellspacing="0" cellpadding="0" style="width:600px"><tbody><tr height="163" style="height:163px"><th style="text-align:right;border-width:medium;border-style:none;border-color:initial;width:140px;vertical-align:top;font-weight:normal;padding:5px;background-color:transparent"><img src="cid:ii_l99s521n0" alt="2013 Dubcovsky UCD NAS Award 2.jpg" width="131" height="155" style="margin-right:0px"><br></th><th style="border-width:medium;border-style:none;border-color:initial;width:440px;vertical-align:top;padding:5px;text-align:left;background-color:transparent"><p align="left" style="margin-bottom:1em;color:rgb(56,56,56);margin-top:0px;line-height:18px;background-color:transparent"><font color="#383838" style="background-color:transparent;font-weight:normal"><font face="tahoma, sans-serif"><b><span style="background-image:initial;background-position:initial;background-size:initial;background-repeat:initial;background-origin:initial;background-clip:initial">El Dr. Jorge Dubcovsky </span></b></font></font><span style="background-color:transparent;font-weight:normal;color:rgb(34,34,34)">es </span><span style="background-color:transparent;font-weight:normal;color:rgb(34,34,34)">Profesor Distinguido en la Universidad de California, Davis, USA, investigador del Howard Hughes Medical Institute y miembro de la Academia de Ciencias de USA. Recibió el premio internacional de Agricultura (“Wolf Prize”) in 2014. El Dr. Dubcovsky ha publicado más de 300 estudios en revistas científicas, que han sido citadas más de 40,000 veces (h= 107), ha liberado 28 cultivares y 72 germoplasmas mejorados, y ha entrenado 29 estudiantes de PhD y 9 de Master. Su laboratorio se enfoca en identificar, entender y utilizar genes de trigo relevantes para el mejoramiento. Su grupo ha desarrollado numerosas herramientas genómicas para acelerar estos descubrimientos, incluidas una colección de mutantes de trigo que incluye 15,000,000 de mutaciones secuenciadas en todos los genes de trigo y sus promotores. El laboratorio del Dr Dubcovsky ha utilizado estas herramientas pare desarrollar variedades de floración temprana y para incrementar el rendimiento y el contenido de proteína y fibra en el grano de trigo.</span></p></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:0px;text-align:left;background-color:rgb(229,229,229)"><table cellspacing="0" cellpadding="0" align="left" style="width:600px"><tbody><tr height="93" style="height:93px"><th style="border-width:medium;border-style:none;border-color:initial;width:570px;vertical-align:middle;padding:10px 10px 0px 20px;text-align:left;background-color:transparent"><p align="left" style="margin-bottom:1em;font-family:verdana,arial,helvetica,sans-serif;color:rgb(56,56,56);margin-top:0px;line-height:18px;background-color:transparent"><span style="font-weight:normal;font-size:14px;line-height:18px"><strong>Certificados:</strong> <span class="gmail_default" style="font-family:tahoma,sans-serif"></span></span><span style="font-weight:normal;background-color:transparent;font-family:Roboto,RobotoDraft,Helvetica,Arial,sans-serif;color:rgb(34,34,34)">los alumnos que necesiten certificar su asistencia, deberán enviar </span><span style="font-weight:normal;background-color:transparent">su nombre y DNI</span><span class="gmail_default" style="font-weight:normal;background-color:transparent;font-family:tahoma,sans-serif"> </span><span class="gmail_default" style="background-color:transparent;font-family:tahoma,sans-serif"><span style="font-weight:normal">al moderador (</span><a href="mailto:mtexeira@agro.uba.ar" style="font-weight:normal" target="_blank">mtexeira@agro.uba.ar</a><span style="font-weight:normal">) </span>durante el transcurso del seminario o en la hora posterior a la finalización del mismo</span><span style="font-weight:normal;background-color:transparent">. </span></p></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:0px;text-align:left;background-color:rgb(66,176,91)"><table cellspacing="0" cellpadding="0" align="left" style="width:600px"><tbody><tr height="62" style="height:62px"><th style="border-width:medium;border-style:none;border-color:initial;width:560px;vertical-align:top;font-weight:normal;padding:25px 20px 10px;text-align:left;background-color:transparent"><p align="center" style="margin-bottom:1em;font-size:12px;font-family:tahoma,geneva,sans-serif;color:rgb(9,9,9);text-align:center;margin-top:0px;line-height:15px;background-color:transparent"><strong><span style="color:rgb(255,255,255)">Instituto de Investigaciones Fisiológicas y Ecológicas Vinculadas a la Agricultura</span></strong></p></th></tr></tbody></table></th></tr><tr><th style="border-width:medium;border-style:none;border-color:initial;width:600px;font-weight:normal;padding:1px 0px;text-align:left;background-color:transparent"><table cellspacing="0" cellpadding="0" align="left" style="width:600px"><tbody><tr height="66" style="height:66px"><th style="border-width:medium;border-style:none;border-color:initial;width:590px;vertical-align:middle;font-weight:normal;padding:5px;text-align:left;background-color:transparent"><p align="center" style="margin-bottom:1em;font-size:12px;font-family:tahoma,geneva,sans-serif;color:rgb(253,251,251);text-align:center;margin-top:0px;line-height:15px;background-color:transparent"><br>IFEVA - Facultad de Agronomía - Universidad de Buenos Aires<br> Av. San Martín 4453 - C1417DSE - Buenos Aires - Argentina</p></th></tr></tbody></table></th></tr></tbody></table></td></tr></tbody></table></div>-- <br><div dir="ltr" data-smartmail="gmail_signature"><div dir="ltr"><div>Marcos Texeira<br>Departamento de Métodos Cuantitativos y Sistemas de Información //  <br>LART - IFEVA - CONICET<br>(011) 452870829</div><div>Facultad de Agronomía<br>Universidad de Buenos Aires</div><div><span>Av. San Martín 4453 (C1417DSE)<br></span></div></div></div></div>
</div></div>